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BLUE-WINGED WARBLER  Vermivora cyanoptera (pinus) 11.5–12cm. Winter

migrant to humid forest and edge, second growth; more varied habitats in

migration. Low to high, often in dead-leaf clusters and vine tangles; joins mixed

flocks. Distinctive, with dark lores, white wingbars on blue-gray wings, white tail

flashes. Hybrids with Golden-winged Warbler rarely reported, show inter­mediate

characters. SOUNDS: High, slightly buzzy tssi, often doubled. STATUS: Uncommon

Sep–Apr on Caribbean slope, mainly below 1500m; rare in winter on Pacific

slope. (Breeds e. N America, winters Mexico to Panama.)

GOLDEN-WINGED WARBLER  Vermivora chrysoptera 11.5–12cm. Winter

migrant to humid forest and edge; more varied habitats in migration, from beach

scrub to semi-open areas with taller trees. Mainly at mid–upper levels, in dead-

leaf clusters and vine tangles; joins mixed flocks. Distinctive, with dark mask and

throat, yellow wingbars; cf. hybrids with Blue-winged Warbler. SOUNDS: High,

slightly buzzy tssi, often doubled; not readily told from Blue-winged call.

STATUS:  Fairly common to uncommon Sep–Apr on both slopes, locally to

2400m, most numerous in foothills; more widespread in migration but rare in

nw. Pacific lowlands. (Breeds e. N America, winters Mexico to nw. S America.)

NORTHERN PARULA  Setophaga americana 10–11cm. Rare winter migrant to varied wooded habitats,

semi-open areas with hedgerows and taller trees, mangroves. Mainly at mid–upper levels, often holds tail

cocked. Told from Tropical Parula by more extensive gray-blue hood with distinct white eye-arcs, less

extensive yellow on underparts; adult male has slaty breast band. Imm. male and adult female resemble

strongly marked imm. female, sometimes with partial breast band. SOUNDS: Sharp stik. STATUS: Rare late

Oct–Apr on both slopes and in Central Valley; mainly below 1200m, with most records from Caribbean

slope. (Breeds e. N America, winters mainly Mexico and Caribbean.)

TROPICAL PARULA  Setophaga pitiayumi 10–11cm. Humid foothill and

highland forest, adjacent semi-open areas with taller trees; especially in areas

with Spanish moss (Tillandsia spp.). Mainly at mid–upper levels; at times joins

mixed flocks. Distinctive, with mostly deep blue upperparts, rich yellow

underparts, weak white wingbar; lacks white eye-arcs of drabber Northern

Parula. Male has more extensive black mask than female, richer orange suffusion

to breast. SOUNDS: Sharp stik, like Northern Parula. Song a variable, high rapid

twittering and chipping, at times ending with 1–2 short trills, 1.5–3 secs.

STATUS: Fairly common to common on both slopes, mainly 900–2400m on

Pacific slope, 600–1800m on Caribbean slope; rarely descends lower in winter.

(Mexico to S America.)

FLAME-THROATED WARBLER  Oreothlypis gutturalis 11.5–12.5cm. Stunning

resident of highland oak forest, adjacent semi-open areas. Mainly at mid–upper

levels, often foraging actively in canopy; joins mixed flocks. Adult male has

flame-orange throat; also note lack of white wingbars; female and imm. average

duller, often with yellowish throat but same unique plumage pattern; juv.

plumage held briefly, much duller. SOUNDS:  Song a rough, slurred, rather

insect-like buzz, ending emphatically, 1–1.5 secs, zzzzzhhh’ every 1–3 secs, at

times preceded by a rapid ringing twitter, chi-chi-chi-chi-chi-zzzzzhhee, and

variations, 1.5–2 secs. Call a high thin sik, at times in short series. STATUS: Fairly

common to common, mainly 1800m to timberline; occasionally wanders down to 1400m. (Costa Rica

to w. Panama.)